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Welcome! Lemmy is organized into topics called “communities” which have different moderators and rules, but overall share a lot of culture and the same users.
Within communities, people can create topical posts and comment on posts or other comments. You can also vote up good content and down poor content, and higher rated content will be shown more prominently. If you’re familiar with Reddit, it’s a similar organizing principle.
There are lots of great apps that are quite diverse, and on multiple platforms. Be sure to check them out here!
On most cases, there is not much difference in experience between different servers/instances. There isn’t really a local timeline (as in eg Mastodon).
Some exceptions: If you post NSFW, some instances don’t allow that. If you are political, some instances might be a better fit than others. Finally, there are some servers that are not widely federated.